Manny and Ellie are expecting Sid wants to be part of the new family However Manny refuses to let Sid get involved, leaving Sid feeling completely shut out and hurt.
Meanwhile Sid discovers three enormous eggs and adopts them. The 3 adorable dinos hatch and accept Sid as their mother. Until Momma Dinosaur (a T-Rex) attacks scooping up the 3 dinos along with Sid and takes them through a crack underground to a mysterious world.
Diego, Manny, Ellie (and Crash & Eddie) follow to the mysterious dinosaur world and work together to rescue Sid.
Luckily they meet Buck, a swashbuckling, slightly crazy one eyed weasel who offers to help them find Sid. Crash and Eddie willingly become Buck’s pupils, becoming his 'wingmen of mayhem'.
Then Ellie goes into labour as dinosaurs attack. Everyone is in danger! Will the herd ever get back safely? Will Diego leave the herd and rediscover his inner predator? Will they ever find Sid?
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